Sara’s Publications


In Press

Rittmeyer, E.N., A. Elmonier, & S. Ruane. 2021 Phylogeography of the Rough Greensnake, Opheodrys aestivus (Squamata: Colubridae), using multilocus Sanger sequence and genomic ddRADseq data. Journal of Herpetology, In press. Accepted_Rittmeyer_et al_2021


Ruane, S. 2021. New data from old specimens. Ichthyology & Herpetology 109:392-396. Ruane_2021_NewData

Bernstein, J.M., J.C. Murphy, H.K. Voris, R.M. Brown, & S. Ruane. 2021. Phylogenetics of Mud Snakes (Squamata: Serpentes: Homalopsidae): A paradox of both undescribed diversity and taxonomic inflation. Molecular Phylogenetics and Evolution 160: 107109. Bernstein et al 2021

Burbrink, F.T., …Ruane, S., … & H. Zaher. 2020. Interrogating the genomes of Squamata (lizards, snakes, and amphisbaenians) with novel machine-learning techniques show no support for deeper traditional morphological relationships.  Systematic Biology Burbrink et al 2020

Burbrink, F.T., Ruane, S., …. & C.J. Raxworthy. 2019. The origins and diversification of the exceptionally rich gemsnakes in Madagascar.  Systematic Biology Burbrink_et_al_2019

Goodman, A.M.,… & S. Ruane. 2019. New Record and Range Expansion of Masticophis lateralis Hallowell, 1853 (Squamata, Colubridae) into Western Baja California Sur, Mexico. Check List Journal 15: 345-348. CheckList_Masticophis

Veerappan, D., Ruane, S., & Gower, D. 2018. A new subfamily of fossorial colubroid snakes from the Western Ghats of peninsular India. Journal of Natural History 52: 2919-2934. A new subfamily of fossorial colubroid snakes from the Western Ghats of peninsular India

Harrington, S.M., deHaan, J.M.*, Shapiro, L.*, & S. Ruane. 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification.  Biological Journal of the Linnean Society 125:61-71. *High school/undergraduate students mentored by S. Ruane Harrington_et_al_2018

Streicher, J. & S. Ruane. 2018. Phylogenomics of snakes. eLS (Wiley Encyclopedia of Life Sciences) [DOI: 10.1002/9780470015902.a0027476] (Invited  Review).

Ruane, S., Richards, S.J., McVay, J.D., Tjaturadi, B., Krey, K., & C.C Austin. 2018. Cryptic and non-cryptic diversity in New Guinea ground snakes of the genus Stegonotus Duméril, Bibron and Duméril, 1854: A description of four new species (Squamata: Colubridae). Journal of Natural  History 52: 13–16, 917–944 (Invited for Special Issue: Integrative Taxonomy of Reptiles). Available here: Cryptic and non cryptic diversity in New Guinea ground snakes


Ruane, S., Myers, E.A., Lo, K.*, Yuen, S.*, Welt, R.S., Juman, M.*, Futterman, I.*, Nussbaum, R.A,  Schneider, G., Burbrink, F.T., & C.J. Raxworthy. 2018. Unrecognized species diversity and new insights into color pattern polymorphism within the widespread Malagasy snake Mimophis (Serpentes: Lamprophiidae). Systematics and Biodiversity. 16: 229-244. *High school student mentored by S.Ruane. Ruane_et_al_2018

Ruane, S., & C.C. Austin. 2017. Phylogenomics using formalin-fixed and 100+ year old intractable natural history specimens. Molecular Ecology Resources, doi: 10.1111/1755-0998.12655 ruane_et_al-2017-molecular_ecology_resources

This 100+ year old snake  Xylophis stenorhynchus was sequenced for thousands of UCEs; Specimen loaned from California Academy of Sciencesxylophis

Burbrink, F.T., Y. Chan, E.A. Myers, S. Ruane, B.T. Smith, & M.J. Hickerson. 2016. Asynchronous demographic responses to Eastern Nearctic Pleistocene climate changes in vertebrate communities. Ecology Letters doi: 10.1111/ele.12695. burbrink_et_al-2016-ecology_letters

Ruane, S., F.T. Burbrink, B. Randriamahatantsoa, & C.J. Raxworthy. 2016. The cat-eyed snakes of Madagascar: Phylogeny and description of a new species of Madagascarophis (Serpentes: Lamprophiidae) from the tsingy of Ankarana. Copeia 104: 712-721. Ruane_et_al_2016_lolo


Ruane, S., C.J. Raxworthy, A.R. Lemmon, E. Lemmon Moriarty, F.T. Burbrink. 2015. Comparing species-tree estimation with large anchored phylogenomic and small Sanger-sequenced molecular datasets: An empirical study on Malagasy pseudoxyrhophiine snakes. BMC Evolutionary Biology 15:221 DOI 10.1186/s12862-015-0503-1 BMC_Pseudoxyrhophiines_Ruane_et_al_2015


Ruane, S., O. Torres-Carvajal, & F.T. Burbrink. 2015. Independent demographic responses to climate change among temperate and tropical milksnakes (Colubridae: Genus Lampropeltis). PLoS ONE, Ruane et al. PLoS ONE 2015

Ruane, S. 2015. Using geometric morphometrics for integrative taxonomy: An examination of head shapes of milksnakes (genus Lampropeltis). Zoological Journal of the Linnean Society DOI: 10.1111/zoj.12245. Ruane_2015

*Ruane, S., R. Bryson, Jr., R.A. Pyron, & F.T. Burbrink. 2014. Coalescent species delimitation in Milksnakes (genus Lampropeltis) and impacts on phylogenetic comparative analyses. Systematic Biology 63:231-250. *Cover image designed by S. Ruane. Ruane_Syst_Biol_2014

Here’s the World Congress of Herpetology 2012 talk I did that accompanies the coalescent species delimitation in milksnakes paper (Winner of SSAR Seibert Award for Systematics, best student presentation).


Myers, E.A., J.A. Rodriguez-Robles, D.F. DeNardo, R.E. Staub, A. Stropoli*, S. Ruane, & F.T. Burbrink. 2013. Multilocus phylogeographic assessment of the California Mountain Kingsnake (Lampropeltis zonata) suggests alternative patterns of diversification for the California Floristic Province. Molecular Ecology 22:5418-5429. *High school student author. Link

Myers, E.A., S. Ruane, K. Knight, J.L. Knight, & J.M. Ray. 2013. Distribution Record of Tantilla alticola Boulenger, 1903 (Squamata: Colubridae) in Coclé Province, Republic of Panama. Check List Journal 9: 151–152. Tantilla_Checklist_2013

Burbrink, F.T., S. Ruane, & R.A. Pyron. 2012. When are adaptive radiations replicated in areas? Ecological opportunity and unexceptional diversification in West Indian Dipsadine snakes (Colubridae: Alsophiini). Journal of Biogeography 39: 465-475. Burbrink_et_al_2012_Alsophiine

Burbrink, F.T., H. Yao*, M. Ingrasci, R. Bryson, T.J. Guiher, & S. Ruane. 2011. Speciation at the Mogollon Rim in the Arizona Mountain Kingsnake (Lampropeltis pyromelana). Molecular Phylogenetics and Evolution 60:445-454. *High school student author. Burbrink_et_al_2011_pyromelana

Ruane, S., R.A. Pyron, & F.T. Burbrink. 2011. Phylogenetic relationships of the Cretaceous frog Beelzebufo from Madagascar and the placement of fossil constraints based on temporal and phylogenetic evidence. Journal of Evolutionary Biology 24: 274-285. Ruane_et_al_2011_Beelzebufo

Ruane, S., S.A. Dinkelacker, & J.B. Iverson. 2008. Demographic and reproductive traits of Blanding’s turtles, Emydoidea blandingii, at the Western edge of the species’ range. Copeia 4:771-779. Ruane_et_al_Copeia_2008

sara ruane alligator snapping

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